Author contributions: R.M. Andrianasolo, S. Hercberg, P. Galan, E. Kesse-Guyot and R. Varraso designed and conducted the research; E. Kesse-Guyot, M. Adjibade, S. Hercberg, P. Galan and R. Varraso provided essential reagents or provided essential materials; R.M. Andrianasolo and R.

 Varraso analysed data or performed statistical analysis; R.M. Andrianasolo, P. Galan and R. Varraso wrote the manuscript and had primary responsibility for final content; R.M. Andrianasolo, E. Kesse-Guyot, M. Adjibade, S. Hercberg, P. Galan, and R. Varraso were involved in interpreting the results and editing the manuscript for important intellectual content; all authors read, edited and approved the final manuscript.